Hamilton's rule.

The kin selection advantage can be quantified by measuring 3 values.

The benefit to the recipient (B)

The cost to the altruist (C)

The coefficient of relatedness (r):


  • Sibling 1 has inherited the allele from parent A.
  • There is a probability of 1/2 that sibling 2 will also inherit this allele from parent A.
  • The r between the two siblings is 1/2 (0.5).

    Kin selection favors altruism when rB > C.